I'm not sure about the others but the Yaesu Fusion radios basically operate at two levels. The digital modes can be used on repeaters and on simplex, producing excellent sound quality. Depending on the radio, you can also send digital messages from one radio to another while talking on the frequency (or so the documentat ion says).
Using a network like Wires-X does require Internet and will not be much use when the big one hits (west coast).
My experience listening to Wires-X on my FTM400XDR was accidental and did not require any reprogramming. I picked it up on two different frequencies that I happened to have preprogrammed and were in my scan set.